Mail won't send in Tiger

We have two Tiger 10.4.2 running G4 and G5 that's connected to the company's Exchange server. For some reason, the Mail.app (no plug-in) on the G5 refuses to send mail to external addresses and internal works with no problem. I've compared the setting on the G4 but noticed no difference.
I've also done the following:
1. Ran Connection Doctor and both in/out connections verified
2. Log-on as a different user but still fail
3. Remove mail.plist file
4. Repair permission
One time, the error message actually said cannot access port 25 and has it got anything to do with the Firewall software? Both G4 and G5 has NetBarrier and I've also tried switching it off but again fail.
Any help? Cheers!

Mick,
Both Macs are set up as Exchange accounts (though tried set up as IMAP and work as well). Not sure what happened cause they've both been working till the occassional glitch, unable to send.
Company's on a Citrix server and the tech guy claim the Mac aren't suppose to use anything else apart from logging on our remote server via Citrix Client and then use OutLook2003. So he's not giving any tech details on the setup.
All I have is the Exchange address and we've set Mail up using that address for Incoming and Outgoing server.

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