Mailing List Replies Come from Wrong Account

Using Mail 2.0.5 under OS X 10.4.3.
I have two mail accounts, let's call them "X" (my main account) and "Y" (an account I use for mailing lists only). I have a Rule that filters all mail to the Y account to one of several mail folders on my mac. This keeps all my mailing list email organized.
Under OS X 10.3, this worked fine. I could read one of these emails, click Reply, and Mail would properly set the sending account to Y.
Under 10.4, Mail insists on setting the account to X. This causes my list mail reply to bounce, and is annoying the heck out of me!
I've tried various combinations of "Send new messages from: ..." but none seem to work. I can't set it to "Y" because I want all new messages to be sent from my main account "X".
Help!

This should not occur.
As an experiment, I have some old saved received messages in a user created mailbox that were addressed to an old email account/address that was cancelled. When selecting Reply for any of these messages, my default Send New Mail from account is chosen as the sending email account/address.
When doing the same for any messages located in a user created mailbox that were addressed to an existing valid email account/address that is not my default Sent New Mail from account selection, the account/email address the message was addressed to is the account chosen as the sending email account/address.
I believe there is some minor corruption to your Mail.app preference file. Let me know if you need the instructions for targeting the preference file which will require re-entering all email account information.

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