Most of my emails have disappeared from mail Inbox

Hello, last night my emails in my Mail inbox disappeared. I didn't notice at first because the last 10 days are still there, but then I realised that 4 years of email from 1 Nov 2012 up to 2008 (around 20,000 messages) are gone. There are a few for 2008 and then the rest - I had emails back to 2005 have also disappeared.
I am not aware of having done anything different last night - all I did was check my mail as usual - and it was only when I tried to search for an email that I realised my inbox now contains only 300 emails. When I go to iCloud it is the same story. Only 303 emails in my inbox. Trash only contains a few hundred, so its like these emails all just disappeared into the ether
I have no idea what to do or how this happened or what might have caused it. I run Lion 10.7.4 on an old black Macbook and have not installed anything new or upgraded software. I don't download unusual attachments so doubt it is a virus.
Is there any way to try to recover them - my inbox is my filing system so years of important stuff is missing. Any help would be very much appreciated.
thanks

We are adepts, but we are not wizards. We're going to need some more information in order to suggest anything.
Version of Aperture and of OS.
More thorough description of what you currently see, and how it differs from what you expect.
Report on any other strangenesses, in either Aperture or elsewhere.
What you were doing when the problem presented itself.
What you have done since then.
The results of that, and how those results differed from what you expected.
Location of your Library.
Location of any referenced Originals.

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Most of my emails have disappeared from mail Inbox

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