Mp4 files (.m4v) from Handbrake won't open in Quicktime.

Suddenly, about a week ago, whenever I try to open these files in QuickTime, I receive a message saying the required codec is missing.  Yet, these same files will open in iTunes.
In Quicktime, when I click on "Find out more," I'm taken to an article titled, "Media formats supported by QuickTime Player," Article: HT3775, which lists MPEG-4 as a supported format in QuickTime.
I do not understand.

You want to convert your DVD video to Quicktime first, and then encode your video with compressor.  Compressor gives you the most options for converting the video.  Apple's compressor gives you lots of options for setting the size and quality of the H.264 file you want to create.  Since it's Apple, the H.264 you create will work in Quicktime.  Compressor is also really cheap at $50 on the Apple store.
There's several good options out there for converting DVD video to Quicktime.  I recommend taking a look at DVDxDV.

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