My iMac's microphone isn't working

whenever i try and facetime or record somthing the microphone isnt picking up anything but i have a look on system preferances and sound and the micrphone is working perfectly but not when i try to do something else! please help as i need to record somthing

Some sound settings are stored in PRAM, so you can try resetting PRAM
http://support.apple.com/kb/HT1379
If that does help, as a test, I would create a new user account in System Preferences Users & Groups pane.  Log out and log in to the new user account.  Be sure to log out instead of using Fast User Switching.
In the new user account, try the same thing with QuickTime Player to record something.  If it works in the new user account, the cause is something in your current user account.  If it still does not work, then the cause the overall system or hardware.  At least that will narrow down the troubleshooting.

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