No audio in safari or other apps

cant hear any audio streaming over the net while in safari, if I get an email attach with a .mov or .wma file cant hear the audio, I have downloaded some things to my hard drive and when playing them in quicktime cant hear audio. if I hit the volume up or down button on keyboard, it makes the beep showing you the adjustment, but cant hear anything out of what im trying to play.
imac intel 17 in Mac OS X (10.4.6)

D,
Most games and browser plugins have a tendency to change your default sound settings resulting in the loss of sound for some applications,
To recover them go to your Applications/Utilities folder and click on Audio/MIDI Setup. Under 'Audio Ouput' make sure your format settings are at 44100.0 Hz. If they are not, simply click on the 'format' pane and choose the appropriate figure.
Alternately, try simply launching GarageBand. Doing this automatically resets the audio back to it's original (44100.0 Hz) default values.
Hope this helps.

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After reviewing your post, it sounds like you cannot receive MMS messages. I would recommend that you read this article, it may be able to help the issue.
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First, try a system reset. It cures many ills and it's quick, easy and harmless...
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