NO AUDIO ON PLAYBACK

Hi everyone,
I'm having a really serious issue. I've got no audio when playing back my files. I've locked tracks. I've muted tracks. I've converted midi files to real instrument files. I've adjusted settings in preferences to maximize for playback. You name it, I've done it.
I'm working on an iMac, OSX 10.4.3 with 2.1 GHz G5, 1.5 GB of SDRAM.
I've trashed any type of large file off my computer and backed them all up on a separate hard drive...
Audio is playing in other applications like iTunes.
Not sure what to do next and I'm in the middle of a fairly large project. Does anyone out there know the magic trick to get this thing working again?
Thanks.

Will you marry me?
I have no idea how or why that preference got messed with, but it did. And that was it...for now at least. Let's see what gremlins creep up next.
Thank you so much. I thought I combed through all the obvious possibilities.

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"iTunes cannot run because it has detected a problem with your audio configuration"
Resolution
QuickTime may need to be reinstalled. Follow the steps below to uninstall QuickTime and then download and install QuickTime using the QuickTime standalone Installer.
On the Start menu, click Control Panel.
Follow the steps appropriate for your Windows operating system below.
Windows XP and Windows 2000: Open the Add or Remove Programs control panel and selectQuickTime in the list of currently installed programs.
Windows Vista and Windows 7: Click Uninstall a program (or if using Windows Vista's Classic View of the Control Panel, click Programs and Features) and select QuickTime in the list of currently installed programs.
Click the Remove button (or Uninstall for Windows Vista and Windows 7) and follow the prompts to remove QuickTime from your computer.
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