No sound on optical audio input but audio signal is there; mid-2007 iMac running OS 10.6.8

I've connected a digital turntable to my iMac via Toslink RCA/optical and I know the cables and converter are working. In the Sound preferences I can see evidence of an audio signal being received but there is no sound. This setup used to work before I had my hard drive replaced.

Erase the drive then install OS X:
Drive Partition and Format
1. Boot from your OS X Installer Disc. After the installer loads select your language and click on the Continue button.  When the menu bar appears select Disk Utility from the Utilities menu.
If you are preparing an external or a non-startup drive, then open Disk Utility in your Utilities folder.
2. After DU loads select your hard drive (this is the entry with the mfgr.'s ID and size) from the left side list. Note the SMART status of the drive in DU's status area.  If it does not say "Verified" then the drive is failing or has failed and will need replacing.  SMART info will not be reported  on external drives. Otherwise, click on the Partition tab in the DU main window.
3. Under the Volume Scheme heading set the number of partitions from the drop down menu to one. Click on the Options button, set the partition scheme to GUID then click on the OK button. Set the format type to Mac OS Extended (Journaled.) Click on the Apply button and wait until the process has completed.
4. Select the volume you just created (this is the sub-entry under the drive entry) from the left side list. Click on the Erase tab in the DU main window.
5. Set the format type to Mac OS Extended (Journaled.) Click on the Security button, check the button for Zero Data and click on OK to return to the Erase window.
6. Click on the Erase button. The format process can take up to several hours depending upon the drive size.
If this doesn't work then the drive is probably bad or some of the new RAM you installed is marginally defective.

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