Only media players play sounds?

The only way I can make sound come out of my MBP is through iTunes or some other media player, like Quicktime or VLC player. No sytem sounds, no iChat sounds, and websites won't make noise in Safari or Firefox.
This happened once before, and rebooting did nothing, but it was cured when firefox crashed (i forget why), but I don't know how to go about forcing that chain of events again.
Macbook Pro 15.4" 2.16 Ghz   Mac OS X (10.4.7)  

I'm sure you've already checkd this, but have you checked to make sure the System Alert sounds in the Sound System Preference is set to an acceptable level?
Also, if you are using some sort of external speakers and/or a headset, verify that your system alert sounds are set to go through the correct speakers.
That would affect System sounds and maybe iChat sounds. It doesn't seem that would be related to websites though. It's worth a shot, however.

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