Only system sounds play from built-in speakers/headphones!

Hi All,
I'm at my wit's end right now. Here is the situation:
I woke up this morning and tried to watch a YouTube video off of facebook- no sound. I opened up iTunes to try and play music- no sound. I opened up iMovie and launched a video clip from my HD- no sound.
I delete a file- barely audible sound. I made an error and got the "bonk" error sound- barely audible. As if I only have it at one ) bar, but its at MAX volume.
So I plugged in headphones- The "empty trash" sound works perfect (loud) at max volume. Error messages sound perfect- but when I try and listen to iTunes/YouTube/ANYTHING... no sound.
Troubleshooting I've done (to no sucess):
Logout/Login with headphones plugged in
Restart with headphones plugged in (both suggested from http://discussions.apple.com/thread.jspa?messageID=12799085&#12799085)
Updated Quicktime (as per http://support.apple.com/kb/HT2178)
SMC Reset
PRAM Reset (http://discussions.apple.com/thread.jspa?messageID=12767451&#12767451)
^ I got VERY excited from the PRAM reset- After self-restarting a second time, the Apple GONG was LOUD (max volume) while all other times it was barely audible.
Unfortunately, it only effected this feature, and nothing else was fixed.
Any suggestions?

So my husband told me that the night before I discovered this issue, he was using his Apple Remote to lower his sound, and it lowered my iMac's sound as well.
He used that same remote to lower, raise, and do anything and everything to my iMac volume, but I haven't had any luck getting my sound back to normal.

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