Replied to icon in mailbox lists

Hi,
I've had this issue before, and been able to fix it, but now I am no longer able to fix it or find what I may have done before
In a nutshell, the little icon in mailbox lists that indicates that a message has been replied to forwarded, etc, and that links to that said reply.. is no longer linking to the reply for me. In other words, it used to be that I could look at my inbox and see that a message had been replied to, and then I could click on that icon to go to the reply. The link no longer works.
I have rebuilt my mailboxes and I have rebuilt the overall Envelope Index file, with no success. Any suggestions?

your Mail preferences may be corrupt. quit Mail and delete (or move to the desktop) the file ~/library/preferences/com.apple.mail.plist. ~ means your home directory. you can get to it by clicking on the house icon in the sidebar of any finder window or by going to /users/username.
then start Mail. you'll have to recreate all your email accounts so make sure you know all their settings. once recreated they will recognize their existing mailboxes. see if Mail behaves after that.
Alternatively, if you have a backup, restore com.apple.mail.plist from backup from before the problems with Mail started. Mail should not be running when you do this.

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