Safari can't connect to the server - still causing problems

Hi
I have read through several topics relating to this problem - but have yet to be able to resolve the issue. So apologies in advance if it has already been answered in one of the many threads.
I have 2 computers both working off the same broadband hub connection.
Both have the same version of the OS and the same version of Safari: OS 10.4.11 and Safari 4.0.4
I have checked that the network settings are the same and all parental control settings are the same.
The laptop will access sites that the desktop will not.
Can anyone shed some light as to why numerous sites I try to access on the iMac result in a Safari message "Safari can’t connect to the server".
Same applies when I attempt the same sites with Firefox.
I have emptied caches, resolved all font issues, addressed terminal and plists etc. etc. - all to no avail.
Help!!!

Adding Open DNS codes to your Network Preferences, should give good results in terms of speed-up as well as added security:
If you are using a single computer: Open System Preferences/Network. Double click on your connection type, or select it in the drop-down menu, and in the box marked 'DNS Servers' add the following two numbers:
208.67.222.222
208.67.220.220
(You can also enter them if you click on Advanced and then DNS)
Sometimes reversing the order of the DNS numbers can be beneficial in cases where there is a long delay before web pages start to load, and then suddenly load at normal speed:
http://support.apple.com/kb/TS2296
If your computer is part of a network: please refer to this page: http://www.opendns.com/start/bestpractices/#yournetwork and follow the advice given.
You could also try these codes as well: 4.2.2.1 & 4.2.2.2
If you use a Router, make sure it has the latest firmware installed.

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