Smtp relaying error - all of a sudden

In the last week or so, Apple Mail.app has started throwing a "relaying denied" message for mail outbound through smtp-server.nyc.rr.com.
I am not in fact relaying -- this is the smtp server specified (no password) for my own ISP. And nothing about my Mail settings has changed since this began happening. However, and this may be a hint, I think Mail.app may have been updated by an Apple security update during this period.
Oddly, choosing "try later" (which saves it to the outbox) and then opening it again and sending it again *always succeeds.* This is an error that's thrown only the first time "send" is hit.
Any ideas? Thanks in advance.
Mac Mini   Mac OS X (10.4.8)  

It might be an authentication issue. Some SMTP servers will only accept mail if there has been an authenticated POP connection from your IP address within the last 15-20 minutes. Maybe they have changed their timeout.
AK

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