Snow Leopard and Lion on external hard drive

I successfully installed Snow Leopard on an external hard drive in one of the partitions.  I am also able to boot from the Snow Leopard on the partition, access the internet, get my email and have successfully installed Evernote. I am also able to access Apple Store and needed items on my laptop.I am using a MacBook Pro as the laptop so it starts up with Snow Leopard from the external drive and I can still access my original info because I used Super Duper to clone my hard drive (running 10.5.11). My question is when I access email and the internet when Snow Leopard is booting does that change the settings on my MacBook Pro.  All was fine for a couple of days but today I was unable to access the Internet or get my email when I booted from 10.5.8 on my laptop. (The external drive was not connected.) I was able to figure out what to do  to get everything working again but I'm not sure I could repeat it.  So, if I boot from an external drive does that change the settings on the laptop? If so is there a standard way to change things back.  I did make screen shots of the settings on my laptop once I got it working again. 
Thanks.
Nancy

Unless you are using the same common home directory for running both versions of the OS (which is possible) then when booting from two distinct volumes, and it doesn't matter where they are, doing something on one does not affect the other.
You have two home directories, one on each volume.  You are using the one on whichever volume you booted from.  So if you defined your mail settings on each exactly the same way (pop, smtp, etc) I would expect them to behave the same way.  Similarly for the network settings.

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