Suddenly Airtunes Remote Speakers Not Available

I've been using Airtunes for quite a long time with a single problem. Now suddenly I was playing music using iTunes v7.1.1 and the playback stopped. Even so I was able to get to web pages and send and receive email. I noticed that my remote speakers showed at the bottom right of the iTunes window but they were italicized- which means they are unavailable. I then tried connecting to my base station with a wireless connection and it came back. Went back to my wired connection and no remote speakers! OK, so I thought it was the router that the base station and my Mac is connected to so I power-cycled the router and I got the remote speakers back. Started playing music again then a while later (a couple of hours) same thing happened. Power-Cycling the router always brought it back but I thought my router might be failing so I installed a brand new one. I was surprised when later iTunes stopped playing to my remote speakers again. Although I had no need before, I mapped UDP port 5353 to my Airport Express and started playing music. After a while the same thing happened- AirTunes just stopped playing. Power-cycling the router always brings it back, temporarily. So it wasn't the port, and it can't be the router. Anyone have an idea what may be going on? I've see that PC users are having this issue and turning off Bonjour fixes it for them, but I'm using a Mac (PPC) running OS 10.4.9. I did try turning off IPv6 but that didn't help- turned it back on. I'm now thinking it's iTunes 7.1.1. I'll try downgrading to an earlier version to see if that resolves the issue. Anyone have any other ideas?

Until today I had exactly the same problem.
Have you changed or upgraded your anti-virus recently? I have a PowerBookPro and a Dell XP PC on the same wireless network, Netgear main modem/router with 3 Airport Expresses each connected to amps/speakers. Suddenly recently the Mac could still see remote speakers via itunes, but the PC could not.
Turned out it was my upgraded antivirus, upgraded from eset nod32 to eset smart security on default settings. The default eset personal firewall settings are "automatic mode" where the sofware monitors all network traffic and imposes its own rules. I changed this to "interactive mode" where any traffic came up with a dialogue box to ask my permission.
Lo and behold, itunes had been asking permission to acces the airports, but been auto-denied. New setting allowed me to give permission and sure enough the speakers now appear in itunes.
Lesson: try fiddling around with settings of anti-virus monitoring software: it worked for me.
Hope this helps.

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