Task Lists Alerts Will Not Send Using AD Groups

SharePoint 2010 (upgraded from 2003 to 2007 to 2010)
Exchange Server 2013
Alerts on Task Lists will not be sent if "Send Alerts To" is determined by AD group rather than being identified by specific user account.
This was working previous to our Exchange servers being upgraded to 2013, but sometime afterwards (we are not sure it was directly after) they stopped working.  Our test server (straight 2010 install) still sends these alerts.  The "you have
successfully created an alert" still works in both environments.
I have tried this on a new Task List, recreated the alerts, restarted the alert service, still no dice.  The service is clearly still running, since it will send an alert to someone who is specified by their user account instead of just part of the
AD group.  It can't be an Anti-virus or filter issue, since the same users can receive an alert when they are added directly to it, but not if they are added through a group.  We have verified the settings in AD (and it's still working on the test
server anyway) and I can't think of what else to check.
( I followed all of the steps in the marked answer for
this post as well)
Could this be something spcific to the 'upgraded' version of SharePoint?  That is the only obvious difference between our Prod and Test servers.  Does anyone have any other ideas?
As always, thanks for any input!
Elli J Project Solutions Specialist Blog: http://projectserverpants.wordpress.com/

Inderjeet: The groups are both AD and Exchange, that terminology may have escaped me in my explanation.  I'm a weird admin because I know little about the standard windows side and only work in Project Server and SharePoint.
The alerts work on our test server for the groups specified, and the "an alert has been created" function works just fine, so it seems like SharePoint is able to determine who is in the list and send to exchange just fine.  It's just the "Alert me"
function for new items or changed items, etc, that's not sending.  Thanks for your help!
Elli J Project Solutions Specialist Blog: http://projectserverpants.wordpress.com/

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