The other day I set up my Airport Extreme to share a 3TB hard drive and it show up fine in the Airport Utility App. That day was great I could access it on both my Mac's. Now though I can't fine the hard drive in that same App and have tried to fix it.

I've already tried disconnecting and reconnecting the hard drive to the Airport Extreme, No luck.
I've tried adding it all over again. No luck.
Why did it even loose the settings in the first place?

It sounds like you just have 2 formatted drives (SSD, HDD) that are not configured as a Fusion drive. If so, you have to manually move files from the SSD to the HDD and visa versa.
If your system would have come configured with a Fusion drive (SSD and HDD 'fused' together) then OS X would maintain file movement from the SSD to the HDD without user intervention.
Look here then scroll down to Introducing Fusion Drive for a brief explanation.

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