This email cannot be delivered to user@domain because the email address is no longer valid

We have Exchange 2010 and Office 2010 clients. All of our users are experiencing problems when trying to send e-mails to a customer domain. After typing the email address in, a message immediately appears saying: This email cannot be delivered to
user@domain because the email address is no longer valid. If we proceed and send the message anyway, we get the NDR shown below. I have tested from a Hotmail account and the recipient is receiving those mails
fine, the problem is for any user at the recipients domain and from any user at our domain, I have tried using OWA and the same problem so I know it is not the cache. Does any one have any ideas as I have been pulling my hair out  ! Thanks.
Delivery has failed to these recipients or groups:
[email protected]
The email address that you entered couldn't be found. Check the address and try resending the message. If the problem continues, please contact your helpdesk.
Diagnostic information for administrators:
Generating server:
xxx-sbs.xxx.local
[email protected]
#550 5.1.1 RESOLVER.ADR.RecipNotFound; not found ##
Original message headers:
Received: from
xxx-sbs.xxx.local ([fe80::4bbd:1665:5716:1dae]) by
 xxx-sbs.xxx.local ([fe80::4bbd:1665:5716:1dae%10]) with mapi id
 14.02.0387.000; Wed, 14 May 2014 15:35:42 +0100
Content-Type: application/ms-tnef; name="winmail.dat"
Content-Transfer-Encoding: binary
From: SENDE (SENDER EMAIL ADDRESS)
To: "RECIPIENT EMAIL ADDRESS
Subject: SCAN ATTACHED
Thread-Topic: SCAN ATTACHED
Thread-Index: Ac9vgcUbgbouRWzgTjS5UqGhPHduZQ==
Date: Wed, 14 May 2014 15:35:41 +0100
Message-ID: <F645248CCF819B4FA674E2F35CC71EE602ADF254@ xxx-sbs.xxx.local>
Accept-Language: en-GB, en-US
Content-Language: en-US
X-MS-Has-Attach: yes
X-MS-TNEF-Correlator: <[email protected]>
MIME-Version: 1.0
X-Originating-IP: [192.168.16.20]
 

Hi,
Do you mean your domain users can’t send messages to a specific domain? And users can send other external domain messages well? Also check whether you can receive external messages or not.
Please check whether this destination domain is listed in the accepted domain of your Exchange server(EMC > Organization Configuration > Hub Transport > Accepted Domains). If it is, please remove it from the list. Additionally, since the issue happens
to all domain users, I suggest we can collect any logs in your Exchange server to ask a question in Microsoft Exchange Server forum for more professional suggestions:
http://social.technet.microsoft.com/Forums/en-US/home?forum=exchange2010
Regards,
Winnie Liang
TechNet Community Support

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