Two computers can play through AExpr, third one can't!

I'm usually pretty good at trouble-shooting stuff like this, but this time I'm totally stumped!
I have three computers connected via ethernet to an SMC Barricade 7004BR router. Also connected to this router is an Airport Base Station (Graphite). In another room, there's an Airport Express connected to my stereo in "join existing network" mode.
On two of the computers*, I can play music from iTunes to the AExpr speakers. When I select the AExpr from the speakers popup at the bottom of the iTunes window, a message - "connecting to remote speakers..." - appears briefly in the "now playing" (or whatever you'd call it) display at the top of the main iTunes window, and then playback switches to the remote speakers.
On the third computer**, this doesn't work. (And of course, this is the machine with the big hard drive containing my iTunes Library!) This machine can apparently "see" the AExpr - it shows up in the speakers popup at the bottom of the iTunes window. But when I try to select it, a separate AirTunes window appears, saying "Connecting to Airport Express..." This new window displays a barber pole progress bar for a few moments, the AirTunes window closes (no error message or anything), and playback continues through the computer's speakers. If I click on the speakers popup, the AExpr is still listed there. Weird, huh?
I've tried to compare the System Preferences > Network settings and the iTunes > Preference settings on all three computers, and I can't find any significant difference. All three are running OSX 10.4.10. One of the computers that can play is running iTunes 7.4.1. The other two - one that can play and one that can't - are running iTunes 7.4.2. I trashed iTunes from the computer that can't play, downloaded a fresh copy from Apple's website, and re-installed it. Still no go.
Any ideas???
* iBook 500 dual-USB, eMac (can play)
** dual-G4 Gigabit (can't play)

Still no go...
Based on another thread I found here, I went to System Preferences > Network, selected the Built-in Ethernet tab, and used "Assist Me" to create a new location, restarted the computer, and no change. Still can't connect to the remote speakers. (Just on this one computer.)
Then I started from my OSX Tiger disc, ran Disc Utility, repaired the volume (no repairs needed), and repaired permissions, and restarted. I got pretty excited at this point, since several of the repairs mentioned iTunes. But it's still not working.
This seems really weird to me. Why do I get a separate window opening, that says "Connecting to Airport Express..." when my other computers just put a little "Connecting to remote speakers..." message in the "now playing" part of the main iTunes window. Has anyone else seen this before?

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