Unable to find replies - why?

Lately this has been getting worse and worse. Clicking on the curved reply icon, many times instead of opening up the reply I get an error like the following instead:
"Mail was unable to find your response to the message “Meeting next week”. You may have deleted the message."
Needless to say the message was not deleted - it is there in the Sent mail folder.
Any idea why this is happening and how to fix it? I've tried the "Rebuild mailbox" option, it didn't help.
Thanks,
doug

Using Mail 4.4 (OSX 10.6.5)
I have been having this same intermittent problem with Mail using my AOL account. There is no rhyme or reason to when it occurs—with or without attachments. Most outgoing emails get saved, but only some vanish. All of my preferences are set to properly save outgoing messages. Recipients do receive the email fine, but I have no record of it, and clicking the "replied" icon presents the "Unable to find your reply...you may have deleted the message" warning.
I was convinced this was an AOL error, but I now see others using other services have the same problem.
Rebuilding mailboxes doesn't fix it, reloading OSX doesn't fix it, and neither does fixing permissions.
Any vanished outgoing emails also do not show up in "sent message" mailboxes on AOL's web-based mail browser, nor are they on my iPhone. They are being sent, then disappearing before they get archived. And only sometimes, with no common features among the ones that vanish. I have no other solution besides making sure to send a copy to myself.
HELP.

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