USB printer on Airport Express via Bonjour: print queue no longer works

This is a variation of similar postings on this forum, but hopefully provides more information to allow others to pinpoint the problem.
Have been using Intel iMac 10.4.8 now upgraded to 10.4.10. Two printers attached: 1 via USB and a 2nd via USB/Bonjour on Airport Express. Both working flawlessly. 2 days ago, an ADSL Internet outage occurred. When this was reestablished an hour or so later, everything was back to normal, EXCEPT that the USB/Bonjour/Airport printer no longer works. When jobs are sent to it, the printer queue stops and a red "!" appears in the dock on the queue. Restarting the queue just brings back the red "!" again.
Have done all the usual things:
1. Checked cables, restarted iMac, restarted printer, reset Airport.
2. Have confirmed the printer works find if moved to a direct USB port. So the problem is not the printer.
3. Have upgraded the Airport firmware to 6.3. Works just fine as a network access point. So the problem is not the Airport itself.
4. Have deleted, added the printer, renamed the printer - none of these things helped (this was suggested by some postings on the forum)
5. Have tried every combination of which item is powered up first (this was suggested by some postings on this forum)
Have confirmed (by checking CUPS Error Log, as well as by Internet Connection Diagnostics) that the problem is that the iMac can no longer connect to the printer when printing on the local host if it is connected via USB/Bonjour/Airport. However, it can find the printer for adding a printer during setup. So Bonjour is working, but the printing is not.
There are lots of similar postings of people having problems with printers previously working, but then stopping working on an Airport Expresss/Extreme. I hope the above scenario, which is a very simple network topology, and the fact that the iMac and printer and Airport all work correctly on their own, is of help. It does look as if the problem is in the Airport Extreme, and was triggered by the WAN outage, and not fixed by printer reinstallation or firmware upgrade.
Looking forward to someone with a clever suggestion...
LeslieB

What did you use to discover the printers on both Mac and PC. Did you use Bonjour for both?

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