What does it mean when there's a _ at the beginning of a username?

Example: _softwareupdate
Thanks.

OS X started using _ on system groups a few releases back; that convention started with 10.5.
I'd start your own thread for the groups question. As for any group limit, haven't seen that one with OS X; if you're hitting some limit with groups or ACLs or a tool you're using, you'll want to post up some details of the version (10.8?), the volume structure used the output of the id command and the ls -ale@ for the ACL.
The local root account has twenty groups, so I'm not sure what you're seeing with that limit.
Alternatively, you can log a bug directly with Apple asking for the limit to be increased.

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What does it mean when there's a _ at the beginning of a username?

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