What happens when I delete files from my HD

OK, my HD is getting quite full, so at some point I am going to have to take stuff off that I don't need and back them up to DVD or something.
I know that in windows you have to defragment your HD all the time, or it gets messy and starts to run slow.
But Mac OSs work differently, I believe they automatically defrag all content greater than 20 MB, so there is never any need to perform this kind of maintenance.
But what if I start deleting files all over the place, some <20MB, inevitably leaving spaces, how does the system deal with this? It sounds potentially hazardous to do this.
Perhaps someone can explain this to me, suppose I delete a 5MB somewhere in the middle of my HD. Does the system rewrite all the files from that point on, going back 5MB in disk space (if you know what I mean) or does it leave a space, making a note that it is there and waits until a suitable files around the same size comes along and stores it there?

STU9000,
Also for redoing your system you might look into Archive and Install. http://support.apple.com/kb/HT1710
About Archive and Install
Available in Mac OS X 10.2 and later, an Archive and Install installation automatically moves existing system files to a folder named Previous System, then installs Mac OS X again. (See tip 3.) You cannot start up your computer using a Previous System folder, nor can you "re-bless" the Previous System folder. (See tip 1.)
You can choose to preserve your user and network settings before installing. This option automatically moves existing users, Home directories, and network settings. This also skips the Setup Assistant after installation.
Note: Because the "Preserve user and network settings" option moves, not copies, existing users, Home directories, and network settings, from the Previous Systems folder to the newly installed System, these items will not exist in the Previous Systems folder after the Archive and Install is complete.
Weston

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