What is this icon, and what does it do?

When you open an email and are in the read mode, at the bottom of the
Screen, right in the middle there is an icon that looks like a box with an arrow
Pointing into it. When I hit that, the piece of mail I am reading disappears.
I can't find the description even in the iOS manual. What is this icon and what
Is it doing to the mail that disappears? Occasionally I can find that email in the
All mail folder.. But not always.. So confusing!

I believe that is the "archive" button. You have a setting for that instead of delete. Do you see the trash can? If not, that is what it is. Here is a link to the iPhone manual and you can see about changing that, but I would have to say it is probably in settings, mail.
http://manuals.info.apple.com/enUS/iPhone_iOS4_UserGuide.pdf

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