Where the **** have my emails gone?

I am getting this message: The message from S Hey <[email protected]> concerning “Re: rain sand and bears” has not been downloaded from the server. You need to take this account online in order to download it.
Which is bull because the email was downloaded and the account is on 24/7.
I use a POP server and have my Mailbox behaviour set to NEVER delete. Anything over a week old has now gone, including business receipts, transaction details etc. They were there two days ago. Please someone tell me they're not deleted?!

This is one of the many bogus error messages in Mail that have absolutely nothing to do with what the problem really is. What the error message really means in the case of Mail 2.x is that the *.emlx file associated with the message isn’t where the index says it should be.
I have no idea what may have caused your messages to disappear from the filesystem but not from the index, though. Have you tried searching for them in Spotlight?
You may also want to try to salvage as many deleted *.emlx files as possible with a data recovery tool such as Data Rescue II, TechTool Pro, or FileSalvage. Of these, the only one I know for sure that can currently recover deleted *.emlx files is FileSalvage. Stop using your computer right now if you want to try that, as anything you do with the computer may cause the deleted files to be overwritten.

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