Which is the better upgrade: EOS 5DS R or EF 400mm f/4 DO IS II USM?

I have the 5D Mark II with a 100-400 mm lens (the old version). As a wildlife photographer, which would be the better upgrade, the EOS 5DS R or EF 400mm f/4 DO IS II USM?

I always assumed lens upgrade first. In this case I was impressed by the new features of the 5DS R because I photograph small birds, often in trees, often at a distance of 30-50 feet, and almost always moving around. I thought the 61 point high density reticular AF might be quick enough to get a photo (that my current set up can't get) and with a file that would allow severe cropping which would make up for the distance.
On the other hand, photographing birds in riparian areas where there is usually poor lighting might suggest the better lens is the answer. 
That's basically why I thought to ask others.

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