Why do I get a 'NO Symbol' during Mac Mini boot?

My 2.4 GHz Macmini4,1 (with the stock 320G HD), model # A1347 running 10.6.8 Mac OS.
Been great for 3+ years since I've owned it... up until today, (Jan. 1st, 2014)
Now the Mac Mini's boot stalls with a "NO" symbol...
     I've run 'Disc Utility from an 'Option Boot' FireWire back/repair/recovery drive on the Mac mini and it boots fine to be able to perform the Disc Utility software operations... so my Mini's logic board is good.
     Disc Utility says the stock drive is good. Also did a permissions repair while running Disc Utility and it came back with no problems reported.
          Any ideas folks?

     There are no external drives connected directly to the Mini in the normal set up. I use an external 2TB drive for my Time Machine B/U's over my Airport Extreme network that's been in use for several years.
     The Mini can 'Option Boot' .... and I get the orignal internal HD icon, & the FireWire repair drive icon, they both show as bootable source(s).
     Selecting FW Drive the boot continues from FireWire... so my logic board/RAM/Ports all work after booting from a re-purposed OEM 320GB MacBook Pro drive, (to a FireWire Recover/Repair drive).
    (I'm thinking the OS needs to be replaced/restored on the internal drive.)

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Why do I get a 'NO Symbol' during Mac Mini boot?

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