Why do my songs and mps disappear with a exclamation mark when I click on them on ITunes?

I have ITunes downloaded to my PC. I also have an IPad and Ipod. Until recently I was able to  play my Mp3s and whatever I had downloaded on ITunes from teleconferences  and sync them without problem on both devices. Lately however, whenever I click on a song (on my PC ITunes) I immediately get an exclamation mark and a window that asks me if I want to "locate file" then the statement that song "original file cannot be located".
Many of these downloaded talks that have disappeared. (They were downloaded legally by the way by (saving link as.. and on ITune go to File, ad file to librairy. I have had them for 2 or 3 years and don't understand why they are rendered unusable all of a sudden. Can someone help me?
Thank you.

There are a number of ways that the files could have been moved, or that could trigger iTunes into looking in the wrong place. Select a broken track, press CTRL+I to Get Info, click No when asked to locate the file and look at the location on the summary tab. Where is the relevant file really located. If you post back both paths I may have a simple solution, otherwise I have a script that can usually find the "missing" files and reconnect iTunes to them.
tt2

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