Why does file picker unexpectadly closed by Windows message keep hanging Firefox 4 when trying to browse destination folder for download?

Whenever I try to download a file, Firefox 4 asks me to choose a destination folder, then when I browse and choose the folder for the download, Firefox just hangs, and I get the file picker unexpectedly closed by Windows message. I end up having to use task manager to close Firefox.

The post I was responding to said,
"Try to boot the computer in Windows Safe mode with network support (press F8 on the boot screen) as a test to see if that helps. "
Somehow, I think that means Windows...
I've tried Firefox in Safe Mode - it makes no difference.

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