Why is Firefox replacing 'space' with '%20' when I try to save a file?

Alright so when I try to save a file from at least two websites, Firefox is converting the spaces in the file name to '%20'. So for example, instead of 'Firefox rules.doc' it's 'Firefox%20rules.doc'.
This is NOT a html error, Internet Explorer doesn't have these problems on the exact same websites, and it's only a recent occurance for Firefox.
EDIT: Have tried re-installing Firefox; diabling add-ons; diabling plugins; updating plug-ins; creating a new profile.

Are you left-clicking the link when that happens or do you use "Save Link as" the right-click context menu?
Does it also happen if you hold down the Alt key and left-click?

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