Why is My Mac Pro So Slow?

I have a Mac Pro, Quad-Core Intel Xeon, 8 cores, 20 GB Memory. OS 10.6.8
The color wheel spins forever when I start the computer, open mac mail, open a web browser and also working in programs. etc. Many months ago I replaced the 750 GB startup disk with a new, 2 TB, Western Digital Caviar drive. Does anyone have any thoughts on what could be cuasing the computer to operate so slowly? I have 745GB available on the startup drive.
Thanks,
Doug

Do you try to boot Safe Mode?
Tried using Apple Disk Utility Repair from another hard drive?
Another suggestion: move all data off boot drive.
1.2TB of data on boot drive is a lot actually if you can avoid doing so.
A 120GB SSD makes an excellent size and performance for OS and apps for most.
Also clean out browser temp files.
just sounds like corrupt directory, corrupt mail is possible also.
And make sure all yoru RAM is showing up properly.
Then attack the issue with Mail which has its own set of issues and troubleshooting.

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