Why must I sometimes turn off WiFi to download or send email?

I have a Droid Razr Maxx, though I don't think it's an Android or even Verizon issue. The problem, which is intermittent, and happens for my friend also on an iPhone, is that sometimes I'm not able to download or send email when connected to a Wifi hot spot. If I turn off WiFi on the phone, it then works fine using the carrier's data plan. Has anyone else experienced this problem and/or know a solution for it?
Thanks in advance!

Out of Office responses are best set up in the webmail account which you access using a browser.
So check there first.
If you set it up in Thunderbird, then you would have had to leave Computer on and Thunderbird running connected to internet all the time.
You would have created a template email to use. This would be in the 'Templates' folder.
You would also have set up a Message Filter to respond to emails in address book using the Template email.
So check in Message filters for a filter that does this.
Tools > Message filters
or
Menu icon > Message filters > Message filters

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