Will Mail delete messages in my online Gmail account?

I have recently set up Mail on my Mac and have it sync'd to get messages from my Gmail account. I am new to this and I haven't tried this before now and Mail has imported over 300 messages from my account! My question is: If I delete these messages, will they be deleted from my mail account when I check them online? I would like to be able to just check my mail quickly rather than logging in every time in Safari...
Thanks for all your help on this!

Crap! Now, THAT's just perfect! >=[ I didn't want my messages to leave my folders on my Gmail account online! I checked it just now. =[ How can I get them back??
**Thanks for your tip. I un-checked that box so that it doesn't remove a copy from the server...**

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