Wireless speed conundrum

I have two computers on a wireless network. A G4 running 10.3.9 is set up as the server, but changing that doesn't affect this problem. The other computer is a G5 running 10.4.8.
Web browsing speed, email download speed, etc., is comparable on both. However, when I try to transfer files from one to the other, there is a amazing difference in speed. Copying from the G5 to the G4 is no problem. Copying from the G4 to the G5 is incredibly slow.
My question is -- which computer do you think is the bottleneck here, and where would you begin to diagnose the problem? I'm at my wit's end, having tried everything I can think of.
TIA for any responses.

I would try to figure out if the problems was the network, cpu, or disk bound.
For the network, I would use iperf (http://dast.nlanr.net/Projects/Iperf/) to do a raw network test from the G4 to the G5 and then another from the G5 to the G4.
You might also consider, if possible, connecting the two hosts with an ethernet cable and repeating the tests..
If on the ethernet you get similar differences in speed, then the problem is more likely the end hosts and not the network in between. If the rates even out a bit so that they are not so skewed, then it could be something to do with the network.
When you are doing the tests, it might be interesting to see how much cpu each box is using, then see how much memory they are using.. see if either is running into any walls there. Use the activity monitor app to help out.

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