Yosemite Mail partially launches after upgrade--two different Macs

I just upgraded my iMac and MacBook Pro to Yosemite and in both cases, now Mail only partially launches. I can access the main menus but almost all of the submenus are grey and the main message viewer and activity viewer are missing (the menu options to show them are also grey).
Common denominators:
• Both machines have aftermarket SSD drives (Trim Enabler is now disabled).
• Both machines have more than one internal drive (Mail is located on the boot volume in the standard directories).
• Both machines have the same email accounts (8-10, mostly IMAP, 2 Gmail accounts, 1 iCloud account).
• Both machines have LOTS of email. Nearly 4GB of IMAP data and another 10GB+ of On My Mac data.
• Both machines use the latest SpamSieve 2.9.17.
• Both were upgraded from the latest Mavericks.
• Both machines are having trouble using Spotlight to find apps (yes, it's selected in the Spotlight prefs).
What I've unsuccessfully tried:
• Reinstalling 10.10 from recovery partition
• Safe-booting
• Holding Shift on Mail startup
• Removing contents of ~/Library/Mail/Bundles (was already empty)
• Disabling all accounts in System Preferences --> Internet Accounts
• Rebuilding Spotlight Index
• Repairing permissions
• Examining the console logs (nothing logged when I launch Mail)
• Examining Activity Monitor for Mail and com/apple.MailServiceAgent (both at 0.0 CPU, no other apps draining CPU or RAM)
Stranger behavior:
I pulled the following files from ~/Library/
     Mail
     Application Support/AddressBook/MailRecents-v4.abcdmr
     Containers/com.apple.corerecents.recentsd
     Containers/com.apple.mail
     Containers/com.apple.MailServiceAgent
After restarting Mail, I am prompted to add an account. I go through that process and it verifies my account credentials. Then the Messages window finally appears. When I click Check Mail, it says I have entered the wrong password (quadruple-checked for accuracy) and won’t check mail. I disabled SSL to compare but the same results.
The weirdest part is that some of the menu items are still grey, most notably the Activity menu option. This is the same behavior I get when I create a new user account and try to set up the email account. Then, after five minutes or so, Mail unexpectedly quits. When I relaunch, I’m back to square one.
The crash report from that looks like:
Application Specific Information:
*** Terminating app due to uncaught exception 'NSInternalInconsistencyException', reason: 'Absolute path passed into -[MFLocalAccount mailboxForRelativePath:isFilesystemPath:create:]: /Volumes/tonic/Library/Mail/V2/Mailboxes/2011 Archive.mbox'
abort() called
terminating with uncaught exception of type NSException
Most bizarre about the above message is that the file path refers to a sharepoint (“tonic”) on my other computer. I’ve imported mail from there in the past.
Not sure what else to try short of a backup, clean installation and migration. Ideas anyone?
Thanks so much!
-Aaron

Hi netaanya,
Welcome to the Apple Support Communities!
Your iPhone 4s does work with Continuity, but only phone calls. Please refer to the attached article for more information.
Connect your iPhone, iPad, iPod touch, and Mac using Continuity - Apple Support
Handoff is supported by the following iOS devices and requires iOS 8. Instant Hotspot requires one of these iPhone or iPad devices with cellular connectivity and iOS 8.1. Instant Hotspot also requires Personal Hotspot service through your carrier.
Phone 5 or later
iPhone 4s (sharing iPhone calls only)
iPad (4th generation), iPad Air, iPad Air 2
iPad mini, iPad mini with Retina display, iPad mini 3
iPod touch (5th generation)
Cheers,
Joe

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