I have both the 250 gb high performance hard drive and a 1 ter byte hard drive.  What happens when the 250 gb hardrive is full?

I have both the 250 gb high performance hard drive and a 1 ter byte hard drive.  What happens when the 250 gb hardrive is full? This on Imac desktop

Never let a drive get more than 80% to 85% full.
AND keep 50-100 GB of free space on a startup drive to allow proper function of virtual memory in all OS X versions from 10.5x to current versions.
10.4x can get by with 20 GB of free space on the boot drive, but more is always better.
I try and keep all boot drives between 50 and 60% of capacity.
Beyond 60% and up to 90% capacity, seek times increase almost exponentially; performance suffers.

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