Mail disappearing when pressing send

When I press send on an email on my iPhone, the email often disappears compeltely. It does not attempt to send, and is not in the Sent Items or Outbox (Outbox is not visible).
This has happened a number of times on accounts which do work, and happened again this morning when I definitely had a full 3G signal. I have my email accounts set up to use only one SMTP server (the o2 server) with all other servers set to off. How can I retrieve missing email and prevent this happening in future?

Hi Popshop. You’re welcome.
Unfourtunately, it didnt' help.
What do you mean? Sure it helped. It was meant to determine the scope of the problem, not to solve it, and now we know that the problem is system-wide, not some kind of data corruption confined to your home folder.
Verify/repair the startup disk (not just permissions), as described here:
The Repair functions of Disk Utility: what's it all about?
If the problem persists after having fixed all filesystem issues, reinstall the Combo Update for the type of computer and the version of Mac OS X you’re using, unless this is the version of Mac OS X that came with your computer:
About the Mac OS X 10.4.8 Combo Update
Mac OS X 10.4.8 Combo Update for PPC
Mac OS X 10.4.8 Combo Update for Intel
Take a look at the following articles for guidelines on how to properly install system updates:
Troubleshooting installation and software updates
Installing software updates
Basically, you should verify/repair the startup disk before installing the update, no applications should be running while installing it, and you may experience unexpected results if you have third-party system software modifications (not normal applications) installed.

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