I have just agreed to software updates on my imac and now can't open my mail.  There is a message saying this version of mail isn't compatible with this latest software.  I have located the updated mail in the applications folder but it won't open. Help!

I have just agreed to software updates on my imac and now can't open my mail.  There is a message saying this version of mail isn't compatible with this latest software.  I have located the updated mail in the applications folder but it won't open. I don't have the original operating software disk.  Does anyone have any suggestions?

Your reply suggests that you have moved Mail to a different location, which (as I said) will prevent it from being properly updated. You cannot use Mail 4.5 with any recent version of Mac OS X. Mac OS X 10.7 (Lion) included Mail 5 and Mac OS X 10.8 (Mountain Lion) includes Mail 6. I'm assuming that you have either Lion or Mountain Lion, but if that is incorrect, you will need to provide that information.
At this point, you will need to reinstall your system. Hold down command-R at startup and reinstall the system right on top of the old one. This should not disturb any of your documents or applications, though you should have backups beforehand in case something goes wrong. And at this point, with your system in an uncertain state (I don't entirely know what you have done to it), something could go wrong.
You will also want to delete the old copy of Mail 4.5, wherever that is. You can't use it.
In the future, do not move any of the preinstalled applications. Leave them where they are.

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