IMac Connects to Internet, but not File Server

Got one iMac (Core 2 Duo, OS X 10.5.8) that has a network issue. It can always connect to the internet, but sometimes it stops seeing a file server on the network (Tiger server). The server connection will drop out on this computer, but not on any of the other computers on the network. Things I've tried that don't work:
- restarting computer
- re-seating ethernet cable
- trying new ethernet cable
- zapping PRAM on computer
The connection to the server just comes and goes randomly, but only on this computer. Suggestions? Thanks!

Same here with an iMac 27" connecting to a windows server.

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