Mail accounts fail after a few hours

iMac 24" OS 10.6.8 / Mail 4.6
A month ago my Mail accounts intermittantly stopped connecting to the incoming and outgoing servers, and as of last week the connections are completely down.
I use two email addresses associated with my own domains; one is a .com and one is a .me. Each address has a POP account in Apple Mail, though of course the incoming and outgoing server settings and ports are the same for both accounts.
When I run Connection Doctor, sometimes I'll get red lights indicating failures, and some times I get a green light on both inbound and outbound connections on both accounts. Whether it's red or green doesn't seem to matter, as the mail doesn't send or receive, no matter the test result.
I was running Sophos, so I uninstalled that and re-booted.
I've re-booted the modem and checked for firmware updates, though the internet connection hasn't ever been a problem.
I deleted the Mail accounts, deleted the Mail plist file, deleted the passwords from Keychain, and re-created the accounts from scratch. Both accounts ran (hello to a month's worth of never-received emails!) for a short while, but within a few hours neither account would connect to the servers. The error messages vary, either "Cannot connect to the server, would you like to try another server?" or "Enter the password..." Neither server ever works, and passwords aren't recognized.
It gets weirder. I have another iMac running 10.8.5, and the .com account operates without fail, while the .me account cannot connect - even though the settings are identical. When I try to send from the .me account I get a dialog box asking me to try the .com account's server, which works every time. Mail flows flawlessly to both accounts on my iPod Touch 4th Gen, and to my MacBook Air.
My host is Godaddy, my ISP is Century Link. Both accounts are fully functional via Godaddy's webmail-based servers. Century Link has no suggestions. Does anyone have a wizard of an idea?

No
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