Mail reply address not recognized by server

On Apple Mail using Road Runner server my replies don't get sent because "the server does not accept my address" as the sender. I can't send new messages either. Incoming mail to the same address works fine. My address is unchanged from when it worked. My device is an Apple Air notebook, new, 11" running 10.9.3 Maverick.

I still haven't found a solution to this problem. I purchased Office for Mac with Outlook 2011 because my IT guy told me that there were lots of people using Macs and that they used this program. It worked for 3 days and then I encountered the exact same problem. See below.
Mail could not be received at this time
The server for account "xxxxx" returned the error "Logon failure: unknown user name or bad password." Your username/password or security settings may be incorrect. Would you like to try re-entering your password?
I've asked my IT department and they have no idea what's going on. Apparently my IP address doesn't even show up on their servers. Can someone please help me?
Thanks.
- Peter

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