Office 365 ability for a specific setup - advice please

I have a client, that has 1 main company with 2 areas HM is the main company HC is the secondary company which is part of HM
All of the staff work for HM and HC, and have roles within both... but to clients have a slightly different outward appearance
I have managed to sell the idea of Office 365 to them as they are a small company (one company)
They have 2 domains HM.com and HC.com all of the work towards the HM division go to HM and from HM, and all of the work to the HC area go to and from HC.
This is where the difficulty sets in.
I have sold the idea of a hosted exchange setup as they have used Exchange in the past and see the immense value... but as a small company and SBS no longer available Office 365 is clearly the way to go.
They will definitely be going for a Office E3 plan at £15 a month per user for Office and Exchange (although I do wonder can we combine E3 and Business Premium as not all users require Access?)
Now this will be to the main company of HM as that is the primary company.
The second division of the company HC do need email too though. Now I know I can tie multiple domains into the 1 Exchange email account
So all emails to HP.com and HC.com would be received the 1 user mailbox
However it is the sending from that is important.... can it be set up so;
1. if they reply to an email received to HC it will be from HC (i.e. so the customers don't see their email to HC coming from HM)
2. If they do a new email can they chose to send from HC instead of HM (HM would be the typical default)
If this set up is possible can anyone forward me to a guide as I would like to test or advise
Or would they need to do an Office 365 E3 for HM
and then a hosted Exchange plan from Microsoft for HC
Any help would b gratefully appreciated
thanks
Tris

Hi Tris,
Thanks for your question. This is the forum to discuss questions and feedback for Microsoft Office client. There is not so much about Office 365
Plans/Services/Deployment aspects here, I would suggest you to post in the dedicated forum of
Office 365 Community, where you can get more experienced responses:
http://community.office365.com/en-us/f/default.aspx
The reason why we recommend posting appropriately is you will get the most qualified pool of respondents, and other partners who read the forums regularly can either share their knowledge or learn from your interaction with us. Thank you for your understanding.
Regards,
Ethan Hua
TechNet Community Support
It's recommended to download and install
Configuration Analyzer Tool (OffCAT), which is developed by Microsoft Support teams. Once the tool is installed, you can run it at any time to scan for hundreds of known issues in Office
programs.

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