ONLY Office365 users sending to our domain getting '550 4.4.7 QUEUE.Expired; message expired' errors

We are having issues with Office365 users sending to our exchange server. We are a small exchange shop. No office365 here. But we have had 3 DIFFERENT domains sending emails our direction receiving the '550 4.4.7 QUEUE.Expired; message expired' returned
email error messages. When using gmail or any other mail service we've tried this issue does not happen. What does Office365/hosted exchange look for differently than any other service?
Regards,
Jay

Diagnostic information for administrators:
Generating server: DM2PR07MB591.namprd07.prod.outlook.com
Total retry attempts: 50
[email protected]
Remote Server returned '550 4.4.7 QUEUE.Expired; message expired'
Original message headers:
Received: from DM2PR07MB528.namprd07.prod.outlook.com (10.141.157.19) by
DM2PR07MB591.namprd07.prod.outlook.com (10.141.176.142) with Microsoft SMTP
Server (TLS) id 15.0.1024.12; Mon, 15 Sep 2014 17:09:01 +0000
Received: from DM2PR07MB528.namprd07.prod.outlook.com ([10.141.157.19]) by
DM2PR07MB528.namprd07.prod.outlook.com ([10.141.157.19]) with mapi id
15.00.1024.012; Mon, 15 Sep 2014 17:09:01 +0000
From: USER [email protected]
To: USER2 [email protected]: subject here
Thread-Topic: subject here
Thread-Index: AQHP0QdOHkZOlFhB10CYHLOZcOgKJA==
Date: Mon, 15 Sep 2014 17:09:00 +0000
Message-ID: [email protected]
Accept-Language: en-US
Content-Language: en-US
X-MS-Has-Attach:
X-MS-TNEF-Correlator:
x-ms-exchange-transport-fromentityheader: Hosted
x-originating-ip: [55.55.55.55]
x-microsoft-antispam: BCL:0;PCL:0;RULEID:;UriScan:;
x-forefront-prvs: 03355EE97E
x-forefront-antispam-report: SFV:NSPM;SFS:(10019020)(6009001)(199003)(189002)(81342001)(90102001)(36756003)(81542001)(4396001)(117636001)(2656002)(97736003)(46102001)(50986999)(85306004)(54356999)(20776003)(16236675004)(99286002)(95666004)(19580395003)(83322001)(19580405001)(99396002)(101416001)(80022001)(66066001)(21056001)(76482001)(64706001)(77982001)(107046002)(105586002)(74502001)(87936001)(19627405001)(86362001)(106116001)(106356001)(74662001)(79102001)(85852003)(83072002)(92566001)(92726001)(229853001)(19625215002)(31966008);DIR:OUT;SFP:1102;SCL:1;SRVR:DM2PR07MB591;H:DM2PR07MB528.namprd07.prod.outlook.com;FPR:;MLV:sfv;PTR:InfoNoRecords;A:1;MX:1;LANG:en;
Content-Type: multipart/alternative;
boundary="_000_141080094032474853domain_"
MIME-Version: 1.0
X-OriginatorOrg: domain.com

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Strange Problem.
I have stored 4 accounts on my iPhone and get mails only from 3 from the same PO.
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Originally Posted by laurabuckley
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