Lost bookmarks saving in older versions

When using Acrobat 9, PDF Optimizer, to save to Acrobat 5.0 or 4.0 formats I lose bookmarks. They are retained if I save to Acrobat 6.0 format.

Update: When "Transparency" is checked in the PDF Optimizer options and flattening of transparent objects takes place, bookmarks are lost, regardless of which version it is optimized to, including Acrobat 9. (My previous comment was incorrect).
When I uncheck this option, bookmarks are retained. When saving to Acrobat 4.0, it indicates that flattening is required, so I lose bookmarks only when the target version is Acrobat 4.
This does not seem to be a problem in Acrobat 8 Professional and I went back to using it to generate my PDF.

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