Unable to edit email accounts mac mail preferences

I make changes - like deleting an account and adding a new one - in Mac Mail preferences, but I don't get the pop up asking if I want to save the changes. So after I turn off Mail and turn it back on, the changes have disappeared. Any way to fix this so I can get rid of my old account and add my new one?
Thanks!

I went to the local Apple store earlier this week and asked on of the folks at the Genius Bar if he knew how to set the outgoing mail address. He showed me that I could enable the "reply to" field and then put my mail forwarding address into that field. Unfortunately, there is no easy way to pre-populate that field so that it always has my mail forwarding account address.

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