How to correct Mail 5.3 errors in OS 10.7.5 update on Mac Pro

Since downloading update to OS10.7.5 from Software Update in December, Mail (5.3) cannot send out and all SMTP servers are listed as "Offline". What has caused this and what's the fix??
I have 3 accounts in Mail, from 3 different mail clients with 3 different SMTP outgoing servers. I cannot send any outgoin mail and one one account cannot even open any INBOX mail listed prior to 29th December when I did the update of OS10.7.5. On the other two accounts I can open previous mail.
Have checked the settings of all 3 accounts and seem ok. I have Entourage on the same machine and there is no problem using the same 3 accounts on MS Office Entourage. So seems that it is Mail that is affected. But why and what's the fix?
I have tried and downloading installing the latest update as Mac OS10.7.5 Combo, but it made no difference? I am using a mid-2011 MacPro 2.3 Ghz Intel Core i5.
Grateful for any suggestions, advice etc.
I have put this problem up last week but have been struck by the number of similar Mail problems reported and the curious shortage of replies, let alone suggestions for fixes. Is there something about Mail problems that are so different form the general range of OS/software  problems in Macs???

Did verify disk permission, repair disk permissions
Steffi, since you can not boot I will assume you did this from your install DVD under Disk Utility. Go back again and this time run the Disk Repair, if there are any error (red) message run the Repair again. Then check the SMART Status at the bottom of the window, it should say "Verified". Quit DU, reboot, then download the 10.5.3 Combo Update from the Apple site and try it again this time without any peripheral attached.
http://www.apple.com/downloads/macosx/apple/macosx_updates/macosx1053comboupdate .html
JG

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