Upstream DNS preventing mail

I cannot get my mail server to send messages out. I get the following error:
Jan 14 08:36:47 mail postfix/smtp[23334]: 4C58B1887C2: to=<[email protected]>, relay=mx4.hrnoc.net[216.120.237.53]:25, delay=5.6, delays=0.06/0.04/5.5/0, dsn=4.0.0, status=deferred (host mx4.hrnoc.net[216.120.237.53] refused to talk to me: 553 Your message was rejected because it was sent from a mailserver without a valid PTR (reverse DNS) record. Please contact your ISP.)
Our test domain, oneillhill.net, was bought from GoDaddy and points to zonedit.com where I've set the dns to include this server. Here's a dig:
dig mail.oneillhill.net
; <<>> DiG 9.4.1-P1 <<>> mail.oneillhill.net
;; global options: printcmd
;; Got answer:
;; ->>HEADER<<- opcode: QUERY, status: NOERROR, id: 29981
;; flags: qr rd ra; QUERY: 1, ANSWER: 1, AUTHORITY: 2, ADDITIONAL: 2
;; QUESTION SECTION:
;mail.oneillhill.net. IN A
;; ANSWER SECTION:
mail.oneillhill.net. 7200 IN A 72.20.143.138
;; AUTHORITY SECTION:
oneillhill.net. 7200 IN NS ns3.zoneedit.com.
oneillhill.net. 7200 IN NS ns2.zoneedit.com.
;; ADDITIONAL SECTION:
ns2.zoneedit.com. 172800 IN A 69.72.158.226
ns3.zoneedit.com. 172800 IN A 72.51.43.223
;; Query time: 304 msec
;; SERVER: 10.0.3.12#53(10.0.3.12)
;; WHEN: Mon Jan 14 08:38:50 2008
;; MSG SIZE rcvd: 133
All seems well and it's resolving thru zoneedit.com to us. Here's a dig on the IP address:
dig -x 72.20.143.138
; <<>> DiG 9.4.1-P1 <<>> -x 72.20.143.138
;; global options: printcmd
;; Got answer:
;; ->>HEADER<<- opcode: QUERY, status: NOERROR, id: 5580
;; flags: qr rd ra; QUERY: 1, ANSWER: 1, AUTHORITY: 2, ADDITIONAL: 2
;; QUESTION SECTION:
;138.143.20.72.in-addr.arpa. IN PTR
;; ANSWER SECTION:
138.143.20.72.in-addr.arpa. 38400 IN PTR mail.oneillhill.net.
;; AUTHORITY SECTION:
143.20.72.in-addr.arpa. 38400 IN NS ns2.phonoscope.com.
143.20.72.in-addr.arpa. 38400 IN NS ns1.phonoscope.com.
;; ADDITIONAL SECTION:
ns1.phonoscope.com. 48154 IN A 205.196.176.90
ns2.phonoscope.com. 48154 IN A 72.20.133.20
;; Query time: 229 msec
;; SERVER: 10.0.3.12#53(10.0.3.12)
;; WHEN: Mon Jan 14 08:41:10 2008
;; MSG SIZE rcvd: 159
Phonoscope is our ISP. When I first got the error mentioned at the top of this message I asked them to add an MX record for this mail server.
We have an existing Xserve running OS X Server 10.4 and Kerio Mail at oneillhill.com with the IP address 72.20.143.134. Our plan is to move to OS X Server 10.5 for mail and abandon the Kerio Mail Server. So for testing we're using the .net domain.
I believe DNS is working fine internal to our offices. But this upstream issue must get resolved before we can thoroughly test the mail service. I'm confident using the 134 address and .com on the Leopard test server would work since it's resolving correctly for our current server setup. We have that domain pointed directly to Phonoscope.

Right now, your IP 72.20.143.138 resolves just fine from the outside.
There is no reason a mail server should reject you because of a missing PTR.
The only thing you have right now is that forward and reverse do not necessarily match as you are pointing both your IPs to mail.oneillhill.net. This however should not be an issue and if a mailserver rejects you because of that, it is their problem.
Does this happen to any destination or just to designimations.com? If you like, send me a test message to servertest080114 -at- topicdesk.com. My server does thorough but proper checks.

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