Receive Connectors for Internal Relays - Before & After 2007-2013 Co-existence

I have just completed the process of migrating from Exchange 2007 Standard SP3 RP10 to Exchange 2013 Standard. I now have Exchange 2013 Standard CU3 installed on two Server 2012 Standard systems. The two new exchange servers are members of a DAG.
With Exchange 2007, I had to include the IP addresses of multi-function devices (MFDs) with scan to e-mail capability in a receive connector in order for the scan to e-mail to work. 
Five of the MFDs were on the same subnet as our Exchange 2007 servers, and one MFD was not. 
It is on the same subnet as our Exchange 2013 servers.
I created new Receive connectors on our Exchange 2013 servers for this purpose. 
The one MFD on the same subnet as the Exchange 2013 servers was mistakenly excluded from a receive connector. 
Much to my surprise, it is able to relay mail.
I removed the IP address of one of the other MFDs to see if the fact that it is on a different subnet than the Exchange 2013 servers made a difference. 
I replicated our domain controllers to help ensure that a lack of AD replication would not be an issue with this test. 
The excluded MFD is still able to relay items.
I have an application on a server.  The application relays messages, but the IP address of the server it is on is not included in any receive connectors.
By searching through  the SMTPreceive log, I found the IP address of the MFDs I did my scan to e-mail tests from in the log. 
I also found the IP address and relayed messages from the application server. 
The log indicates that the Default Frontend connector is handling this traffic. 
Does Exchange 2013, by default, relay messages from all internal IP addresses? 
Does this mean that I don’t need receive connectors for any internal devices (MFDs, servers, etc.). 
Our Exchange 2007 servers have been decommissioned.  Would I have needed connectors while we were still in a co-existence situation? 
Am I likely to need them when we have our next co-existence situation in a few years?

Hi Cara. Thank you for the response.
If I understand you correctly, you are saying that the MFDs are able to relay because the Default Frontend connector allows them to. 
That makes sense for the one MFD that was not included in the customised connector I created. 
Thank you for helping me sort that out.
By analysing the Front End Transport logs at the following location on my server, I determined that the included MFDs were using the customised connector I had created. 
C:\Program Files\Microsoft\Exchange Server\V15\TransportRoles\Logs\FrontEnd\ProtocolLog\SmtpReceive
If I removed the IP address for one of those MFDs, it was still able to relay, but it was doing that through the Default Frontend connector. 
That makes sense.  As explained near the end of the following article, Exchange uses the most specific connector. 
http://exchangeserverpro.com/exchange-2013-configure-smtp-relay-connector/
The one that includes the specific IP address of the MFD will be used, but if the IP address is not specified in any connectors, The Default Frontend connector will automatically be used by
the device.
I believe that I now understand how the have the relaying from MFDs is taking place, but I am not certain how some applications are completing their relaying. 
Another department in my organisation has their own network. 
I have a customised receive connector that includes the IP addresses of two application servers on their network. 
Applications on these servers relay messages through the Exchange server on my network. 
The headers on the relayed messages suggest that the messages are being relayed by the Default Frontend connector. 
The logs for that connector, however, do not contain any entries for the two IP address. 
The message tracking logs suggest that the Default <servername> connector is relaying for these applications.
  If the most specific connector is supposed to handle the relaying, why is that not happening in this case? 
Also, two other application servers from this network are not included in any customised connectors I have created. 
The logs show that they are relaying mail through the Default Frontend connector. 
If I understand you correctly, you are saying that this should not be happening, as applications and external users require a customised connector, rather than a built-in connector. 
If I have mis-understood you, please clarify this for me. 
Thanks!!

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    OK so I found some more details online and decided to take the plunge (after a backup was taken) and my problem is now fixed. Although thank you to the 40 people that atleast looked at my query.
    This worked for me, please read, backup and decide yourself if you wish to follow my steps.
    1.  I Read this to understand more on how I broke it in the first place  :
    First section of this......
    https://exchangemaster.wordpress.com/tag/smtp/
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    RemoteIPRanges   : {::-ffff:ffff:ffff:ffff:ffff:ffff:ffff:ffff, 0.0.0.0-255.255.255.255}
    TransportRole    : HubTransport
    PermissionGroups : ExchangeUsers, ExchangeServers
    MaxMessageSize   : 35 MB (36,700,160 bytes)
    Name             : Default Frontend EX2013
    AuthMechanism    : Tls, Integrated, BasicAuth, BasicAuthRequireTLS, ExchangeServer
    RemoteIPRanges   : {::-ffff:ffff:ffff:ffff:ffff:ffff:ffff:ffff, 0.0.0.0-255.255.255.255}
    TransportRole    : FrontendTransport
    PermissionGroups : AnonymousUsers, ExchangeServers, ExchangeLegacyServers
    MaxMessageSize   : 36 MB (37,748,736 bytes)
    Name             : Outbound Proxy Frontend EX2013
    AuthMechanism    : Tls, Integrated, BasicAuth, BasicAuthRequireTLS, ExchangeServer
    RemoteIPRanges   : {::-ffff:ffff:ffff:ffff:ffff:ffff:ffff:ffff, 0.0.0.0-255.255.255.255}
    TransportRole    : FrontendTransport
    PermissionGroups : AnonymousUsers, ExchangeServers
    MaxMessageSize   : 36 MB (37,748,736 bytes)
    Name             : Client Frontend EX2013
    AuthMechanism    : Tls, Integrated, BasicAuth, BasicAuthRequireTLS
    RemoteIPRanges   : {::-ffff:ffff:ffff:ffff:ffff:ffff:ffff:ffff, 0.0.0.0-255.255.255.255}
    TransportRole    : FrontendTransport
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    MaxMessageSize   : 35 MB (36,700,160 bytes)
    4. Recreated the Receive connector for my photocopier/scanner but this time choose Frontend Transport role and Not the default Hubtransport. Restarted the server, crossed my fingers and everything worked!! (Apparently restarting both transport services
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    Source: http://www.msexchange.org/kbase/ExchangeServerTips/ExchangeServer2013/ManagementAdministration/exchange-server-2013-sp1-transport-service-stops-and-does-not-restart.html

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