Connection on default ports timed out

I administer an xserve that up until 3 days ago was sending and receiving email fine. After a user went over quota I adjusted their quota to allow them to remove emails as they were unable to modify any email at all. In the process of adjusting their quota I have broken something and now get "Connection on default ports timed out" error.
I am able to send using webmail but unable to receive from an external source
I have posted my post configuration below and would appreciate any help.
command_directory = /usr/sbin
config_directory = /etc/postfix
content_filter = smtp-amavis:[127.0.0.1]:10024
daemon_directory = /usr/libexec/postfix
debugpeerlevel = 2
enableserveroptions = yes
html_directory = no
inet_interfaces = localhost
mail_owner = _postfix
mailboxsizelimit = 0
mailbox_transport = cyrus
mailq_path = /usr/bin/mailq
manpage_directory = /usr/share/man
messagesizelimit = 26214400
mydestination = $myhostname,localhost.$mydomain,wcc.school.nz
mydomain = wcc.school.nz
mydomain_fallback = localhost
myhostname = mail.wcc.school.nz
mynetworks = 127.0.0.0/8
newaliases_path = /usr/bin/newaliases
queue_directory = /private/var/spool/postfix
readme_directory = /usr/share/doc/postfix
sample_directory = /usr/share/doc/postfix/examples
sendmail_path = /usr/sbin/sendmail
setgid_group = _postdrop
smtpdenforcetls = no
smtpdpw_server_securityoptions = plain,login
smtpdrecipientrestrictions =
permitsasl_authenticated,permit_mynetworks,reject_unauthdestination,permit
smtpdsasl_authenable = yes
smtpdtls_certfile = /etc/certificates/Default.crt
smtpdtls_keyfile = /etc/certificates/Default.key
smtpdtlsloglevel = 0
smtpduse_pwserver = yes
smtpdusetls = no
unknownlocal_recipient_rejectcode = 550

It's not surprising.
Your configuration includes:
inet_interfaces = localhost
This tells postfix to listen on localhost only. i.e. it won't listen on the network interfaces and accept mail from remote clients.
There may be other issues with the configuration, too, but that's the first one to fix.

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